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5'-nucleotidase beta-carotene isomerase 9-cis-beta- carotene 9',10'-cleavage dioxygenase carlactone synthase cytochrome P450 monooxygenase cytochrome P450 monooxygenase Phospholipase D1 B-Carotene 9-cis-β- Carotene O2 9-cis-10'- apo-β- carotenal β-Ionone O2 carlactone (2E,4E,6E)-7- hydroxy-4- methylhepta- 2,4,6-trienal O2 carlactone carboxylate H+ H+ 5-deoxystrigol Iron Fe2+ Heme Heme oxidized electron acceptor reduced electron acceptor oxidized electron acceptor reduced electron acceptor PC Phosphatidic Acid beta-carotene biosynthesis Fe2+ Chloroplast Stroma