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Aldophosphamide Transporter Cytochrome P450 2A6 Cytochrome P450 2B6 Cytochrome P450 2C19 Cytochrome P450 2C8 Cytochrome P450 3A4 Cytochrome P450 3A4 Cytochrome P450 3A4 Retinal dehydrogenase 1 Retinal dehydrogenase 1 Phosphoramide mustard Adenine Aldophosphamide Aldophosphamide Aldophosphamide Phosphoramide mustard Cyclophosphamide Cyclophosphamide Cyclophosphamide O2 NADPH H+ 4-Hydroxycyclophosphamide H2O NADP NAD 4-Ketocyclophosphamide NADH O2 H2O Chloroacetaldehyde 3-Dechloroethylifosfamide H2O2 Acrolein NAD H2O Carboxyphosphamide NADH H+ NAD H2O Acrylic acid NADH H+ Nornitrogen mustard Phosphoramide Aziridinium Cl- Adenosine phosphoester adduct Heme Heme Heme Heme Heme Heme Cytochrome P450 2C9 Aldehyde dehydrogenase, dimeric NADP-preferring Aldehyde dehydrogenase, dimeric NADP-preferring Heme Heme Cyclophosphamide is absorbed in the GI tract and is then metabolized in the liver through various P450 enzymes. Aldophosphamide is is an important metabolite from the breakdown. Aldophosphamide is transported out of the liver cell into the blood via golgi apparatus vesicles. Aldophosphamide is taken up by into cancer cells and then converted to acrolein, phosphoramide mustard, and carbodyphosphamide. Phosphoramide mustard is the main cytotoxic metabolic agent from cyclophosphamide. It moves into the nucleus where it does 3 actions to inhibit DNA replication. Firstly, phosphoramide mustard crosslinks DNA causing it to be unable to separate the strands for replication. Second, phosphoramide induces the mispairing of nucleotide bases during replication causing mutations. This creates fraudulent DNA that cannot properly transcribe RNA. Third, phosphoramide attachs alkyl groups to DNA bases, such as adenine in this reaction, which results in the DNA becoming fragmented due to repair enzymes trying to replace the alkylated bases. This preventing DNA synthesis and RNA transcription from affected DNA that has become alkylated. Hepatocyte Apoptosis The miscoding of the DNA, the cross-linking of guanine bases and the fragmentation of the DNA all lead to the apoptosis of the cancerous cell. Cancerous Cell Nucleus Liver
Nucleus Nucleus Unknown CYP2A6 CYP2B6 CYP2C19 CYP2C8 CYP3A4 CYP3A4 CYP3A4 ALDH1A1 ALDH1A1 Phosphoramide mustard Adenine Aldophosphamide Aldophosphamide Aldophosphamide Phosphoramide mustard Cyclophosphamide Cyclophosphamide Cyclophosphamide Oxygen NADPH Hydrogen Ion 4- Hydroxycyclophosphamide Water NADP NAD 4- Ketocyclophosphamide NADH Oxygen Water Chloroacetaldehyde 3- Dechloroethylifosfamide Hydrogen peroxide Acrolein NAD Water Carboxyphosphamide NADH Hydrogen Ion NAD Water Acrylic acid NADH Hydrogen Ion Nornitrogen mustard Phosphoramide Aziridinium Chloride ion Adenosine phosphoester adduct CYP2C9 ALDH3A1 ALDH3A1
CYP2A6 CYP2B6 CYP2C19 CYP2C8 CYP3A4 CYP3A4 CYP3A4 ALDH1A1 ALDH1A1 Phospho Adenine Aldoph Aldoph Aldoph Phospho Cycloph Cycloph Cycloph O2 NADPH H+ 4HCP H2O NADP NAD 4KcycPm NADH O2 H2O chlogl 3DCLFA H2O2 Acrolen NAD H2O Caroxyp NADH H+ NAD H2O AcryliA NADH H+ Nornm PhoAzdm Cl- Ad ph a Heme Heme Heme Heme Heme Heme CYP2C9 ALDH3A1 ALDH3A1 Heme Heme Cyclophosphamide is absorbed in the GI tract and is then metabolized in the liver through various P450 enzymes. Aldophosphamide is is an important metabolite from the breakdown. Aldophosphamide is transported out of the liver cell into the blood via golgi apparatus vesicles. Aldophosphamide is taken up by into cancer cells and then converted to acrolein, phosphoramide mustard, and carbodyphosphamide. Phosphoramide mustard is the main cytotoxic metabolic agent from cyclophosphamide. It moves into the nucleus where it does 3 actions to inhibit DNA replication. Firstly, phosphoramide mustard crosslinks DNA causing it to be unable to separate the strands for replication. Second, phosphoramide induces the mispairing of nucleotide bases during replication causing mutations. This creates fraudulent DNA that cannot properly transcribe RNA. Third, phosphoramide attachs alkyl groups to DNA bases, such as adenine in this reaction, which results in the DNA becoming fragmented due to repair enzymes trying to replace the alkylated bases. This preventing DNA synthesis and RNA transcription from affected DNA that has become alkylated. Hepatocyte Apoptosis The miscoding of the DNA, the cross-linking of guanine bases and the fragmentation of the DNA all lead to the apoptosis of the cancerous cell. Cancerous Cell Nucleus Liver
Nucleus Nucleus CYP2A6 CYP2B6 CYP2C19 CYP2C8 CYP3A4 CYP3A4 CYP3A4 ALDH1A1 ALDH1A1 Phospho Adenine Aldoph Aldoph Aldoph Phospho Cycloph Cycloph Cycloph O2 NADPH H+ 4HCP H2O NADP NAD 4KcycPm NADH O2 H2O chlogl 3DCLFA H2O2 Acrolen NAD H2O Caroxyp NADH H+ NAD H2O AcryliA NADH H+ Nornm PhoAzdm Cl Ad ph a CYP2C9 ALDH3A1 ALDH3A1